PATRICK D. MOLDOWAN
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Peer-reviewed publications
In review/revision
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​Giacometti D,
 PD Moldowan, GJ Tattersall. Sub-zero temperatures during early spring migration in Blue-spotted salamanders (Ambystoma laterale).


Moldowan, PD.
 Hyperpredation of freshwater turtles and tortoises by subsidized corvids: global case studies of a conservation conundrum. (... preview)


2022
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​Moldowan PD, 
GJ Tattersall, N Rollinson. ​2022. Climate-associated decline of body condition in a fossorial salamander. Global Change Biology 28: 1725–1739. [PDF]
2020
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Moldowan PD, RJ Brooks, and JD Litzgus. 2020. Sex, shells, and weaponry: Coercive reproductive tactics in the Painted Turtle, Chrysemys picta. Behavioral Ecology and Sociobiology 74: 142. [PDF]


Terebiznik* M., PD Moldowan*, JA Leivesley, MD Massey, C Lacroix, JWH Connoy, N Rollinson. 2020. Hatchling turtles ingest natural and artificial incubation substrates at high frequency. Behavioral Ecology and Sociobiology. 74: 130. [PDF] *equal contributions

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​Moldowan PD.
 Pseudemys concinna (River Cooter). Wounding, conspecific aggression, reproductive behavior. Herpetological Review 51(3): 582–583. [PDF]


​Moldowan PD, RJ Brooks, and JD Litzgus. 2020. Demographics of injuries indicate sexual coercion in a population of Painted Turtle (Chrysemys picta). Canadian Journal of Zoology 98(4): 269–278. [PDF]
​Press: Herpetological Highlights.
2019
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Moldowan PD. 2019. Lizard pollination, or the stigma of having food on your face. Frontiers In Ecology and the Environment 17(10): 574. [PDF]

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Hawkshaw* DM, PD Moldowan*, RJ Brooks, JD Litzgus, and N Rollinson. 2019. Discovery and description of a novel sexual weapon in the world’s most widely-studied freshwater turtle. Evolutionary Ecology 33(6): 889–900. *equal contributions. [PDF, also see correction] 
Press: New York Times.

Moldowan* PD, MA Smith*, T Baldwin, T. Bartley, N Rollinson, and H Wynen. 2019. Nature’s pitfall trap: Salamanders as rich prey for carnivorous plants in a nutrient-poor northern bog ecosystem. Ecology 100(10): e02770. [PDF]  *equal contributions
Examples of press: National Geographic, Smithsonian Magazine, CBC, New Scientist, Live Science, University of Toronto, In Defense of Plants, Mother Nature Network, Herpetological Highlights

Francis EA, PD Moldowan, MA Greischar, and N Rollinson. 2019. Anthropogenic nest sites provide warmer incubation environments than natural nest sites in a population of oviparous reptiles near their northern range limit. Oecologica 190(3): 511–522. [PDF]
2018
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Moldowan PD, RJ Brooks, and JD Litzgus. 2018. Sex-biased seasonal capture rates in the Painted Turtle, Chrysemys picta. Canadian Field-Naturalist 132(1): 20–24. [PDF]
2017
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Moldowan PD, RJ Brooks, and JD Litzgus. 2017. Assessing head morphology dimorphism in the Midland Painted Turtle (Chrysemys picta marginata) using a photographic questionnaire. Chelonian Conservation and Biology 16(1): 76–82. [PDF]
2016
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Moldowan PD 
and H. Kitching. 2016. Observation of an Eastern Wolf (Canis sp. cf. lycaon) caching food in a Sphagnum bog in Algonquin Provincial Park, Ontario, Canada. Canadian Field-Naturalist 130(4): 351–354. [PDF]
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Moldowan PD, TM Laverty, CJ Emmans, and RC Stanley. 2016. Diet, gastric parasitism, and injuries of caimans (
Caiman, Melanosuchus, and Paleosuchus) in the Peruvian Amazon. South American Journal of Herpetology 11(3): 176–182. [PDF]


​Moldowan PD, RJ Brooks, and JD Litzgus. 2016. Quantification of cranial and tomiodont dimorphism in Testudines using the Midland Painted Turtle, Chrysemys picta marginata. Zoomorphology 135(4): 499–510. [PDF]

Moldowan PD. 2016. Bothrops atrox (Common Lancehead). Diet and fang replacement. Herpetological Review 47(3): 473–474. [PDF]

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​Moldowan PD
, JA Copsey, N Zuël, V Tatayah, and N Cole. 2016. Sticks and stones: notes on the ecology and conservation of an endemic stick insect (Apterograeffea marshallae) and the restoration of an island ecosystem. Phelsuma 24(2016): 72–79. [PDF]

Moldowan PD, RJ Brooks, and JD Litzgus..  2016. Turtles with “teeth”: beak morphology of Testudines with a focus on the tomiodonts of Painted Turtles (Chrysemys ​spp.). Zoomorphology 135(1): 121–135. [PDF]
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2015
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Moldowan PD, MG Keevil, PB Mills, RJ Brooks, and JD Litzgus. 2015. Diet and feeding behaviour of Snapping Turtles (Chelydra serpentina) and Midland Painted Turtles (Chrysemys picta marginata) in Algonquin Provincial Park, Ontario. Canadian Field-Naturalist 29(4):403–408. [PDF]

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Moldowan, PD, MG Keevil, N Koper, RJ Brooks, and JD Litzgus. 2015. Growth and sexual maturity of a female Midland Painted Turtle (Chrysemys picta marginata) afflicted with kyphosis. Chelonian Conservation and Biology 14: 157–160. [PDF]

​Moldowan PD, MG Keevil, S Kell, RJ Brooks, and JD Litzgus. 2015. Anti-predator defences of Brown Bullheads (Ameiurus nebulosus) and interactions with Snapping Turtles (Chelydra serpentina). Canadian Field-Naturalist 129(2): 189–193. [PDF]


​Bobbie CB, PD Moldowan, and BA Bobbie. 2015. Chelydra serpentina (Snapping Turtle). Waterfowl predation. Herpetological Review 46(1): 77. [PDF, see page 77]


Baxter-Gilbert J, JL Riley, PD Moldowan, and JD Litzgus. 2015. Sistrurus catenatus catenatus (Eastern Massasauga) and Nerodia sipedon sipedon (Northern Watersnake). Diet, road mortality, and foreign objects. Herpetological Review 46(1): 107. [PDF, see page 107]
2013
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Moldowan PD, DL LeGros., and GJ Tattersall. 2013. Lithobates sylvaticus (Wood Frog). Davian Behaviour. Herpetological Review 44(2): 297. [PDF]

Boyle SP and PD Moldowan. 2013. Ambystoma maculatum (Spotted Salamander). Terrestrial oviposition. Herpetological Review 44(1): 113–114. [PDF]

2011
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Riley JL, M Keevil., P Moldowan, and JD Litzgus. 2011. Emydoidea blandingii (Blanding’s Turtle). Record egg size. Herpetological Review 42(3): 417–418. [PDF]

Book Contributions
  • Species Conservation: Lessons from Islands (2018) by JA Copsey, SA Black, JJ Groombridge and CG Jones (written)
  • Mammals of Algonquin Provincial Park (2018) by D. Strickland, R. Rutter and S. Dobbyn (photographic)
  • Wildlife on Roads, a Handbook (2018) by K.E. Gunson and F.W. Schueler (editorial and photographic)
  • North American Amphibians, Distribution and Diversity (2013) by D.A. Green, L.A. Weir, G.S. Casper, and M. Lannoo (photographic)
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"Popular" Writing
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Recent sampling ...
  • "On Island Time, an island-hopping sabbatical-of-sorts", The EEB Quarterly (2021)
  • ​"Canadian Herpetologists Abroad: New Zealand for the World Congress of Herpetology" for the Canadian Herpetological Society (2020)
  • "A Nod to the Lava Lizards of the Galapagos", for Quest Nature Tours (2020)
  • ​"Evolution Gone Wild: A Reflection on Darwin’s Enchanted Isles", for Quest Nature Tours (2020)
  • "75 Years of Wildlife Research", for Algonquin Provincial Park's The Raven (2019)


Art meets science: Fortuitous photographs and serendipitous inspiration (2018; on Robert Bateman and Robert MacArthur at the Algonquin Wildlife Research Station). EEB Quarterly 1(1): 70-77. <Web Link> 

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Antler flies and the moose antler - an ecosystem unto itself! 2016. Ontario Nature blog.


Crown jewels in Canadian herpetology and herpetoculture: a tribute to Karel Fortyn, his Orinoco Crocodiles, and the Seaway Serpentarium. 2016. The Canadian Herpetologist 6(1): 7-14. [PDF]

Canada's New Noah blog series on wildlife research and conservation in Mauritius, via Wildlife Preservation Canada.  2015.
(1) Beginning the Canada's New Noah experience; (2) Reptile confessions: long tails (tales) and sticky toes;
(3) Even giants start small: life the Aldabra Giant Tortoise;
(4) The fruit bat motto: "Got flowers? Will pollinate! Go seeds? Will disperse!"
(5) A conservation epiphany: Importance of an individual

Secret love life of the Painted Turtle. 2015. The Raven, a natural and cultural history digest. Algonquin Provincial Park, Ontario, Canada.  [PDF]

'Twas the night before field season. 2015. The Canadian Herpetologist 5(1): 6-7. [PDF]


Harbinger of Spring, the Spotted Salamander. 2014, revised 2018. Ontario Parks blog <Web Link 1> <Web Link 2> 


Invited Reviews for:
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Site updated: August 2022
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